The Bee Hummingbird is the smallest bird in the world, weighing 1.8 g (0.06 oz) and measuring about 5 cm (2 in). A typical North American hummingbird, such as the Rufous Hummingbird (Selasphorus rufus), weighs approximately 3 g (0.106 ounces) and has a length of 10–12 cm (3.5–4 inches). The largest hummingbird is the Giant Hummingbird (Patagona gigas), with some weighing as much as 24 g (0.85 oz) and measuring 21.5 cm (8.5 in).

Most species exhibit conspicuous sexual dimorphism, with males brightly colored and females displaying cryptic coloration.Iridescent plumage is present in both sexes of most species, with green being the most common color. Highly modified structures within certain feathers, usually concentrated on the head and breast, produce intense metallic iridescence.

Hummingbirds feed on the nectar of plants and are important pollinators, especially of deep-throated, tubular flowers. Like bees, they are able to assess the amount of sugar in the nectar they eat; they reject flower types that produce nectar which is less than 15% sugar and prefer those whose sugar content is around 25%. Nectar is a poor source of nutrients, so hummingbirds meet their needs for protein, amino acids, vitamins, minerals, etc. by preying on insects and spiders, especially when feeding young.

 

 

Most hummingbirds have bills that are long and straight or nearly so, but in some species the bill shape is adapted for specialized feeding. Thornbills have short, sharp bills adapted for feeding from flowers with short corollas and piercing the bases of longer ones. The Sicklebills’ extremely decurved bills are adapted to extracting nectar from the curved corollas of flowers in the family Gesneriaceae. The bill of the Fiery-tailed Awlbill has an upturned tip, as in the Avocets. The male Tooth-billed Hummingbird has barracuda-like spikes at the tip of its long, straight bill.

The two halves of a hummingbird’s bill have a pronounced overlap, with the lower half (mandible) fitting tightly inside the upper half (maxilla). When hummingbirds feed on nectar, the bill is usually only opened slightly, allowing the tongue to dart out and into the interior of flowers.

Like the similar nectar-feeding sunbirds and unlike other birds, hummingbirds drink by using protrusible grooved or trough-like tongues.

Hummingbirds do not spend all day flying, as the energy costs of this would be prohibitive; the majority of their activity consists simply of sitting or perching. Hummingbirds feed in many small meals, consuming many small invertebrates and up to five times their own body weight in nectar each day. They spend an average of 10-15% of their time feeding and 75-80% sitting and digesting.

 

 

 

Hummingbird flight has been studied intensively from an aerodynamic perspective using wind tunnels and high-speed video cameras.

Writing in Nature, the biomechanist Douglas Warrick and coworkers studied the Rufous Hummingbird, Selasphorus rufus, in a wind tunnel using particle image velocimetry techniques and investigated the lift generated on the bird’s upstroke and downstroke.

They concluded that their subjects produced 75% of their weight support during the down-stroke and 25% during the up-stroke: many earlier studies had assumed (implicitly or explicitly) that lift was generated equally during the two phases of the wingbeat cycle, as is the case of insects of a similar size. This finding shows that hummingbirds’ hovering is similar to, but distinct from, that of hovering insects such as the hawk moths.

The Giant Hummingbird’s wings beat at 8–10 beats per second, the wings of medium-sized hummingbirds beat about 20–25 beats per second and the smallest beat 70 beats per second.

 

 

As far as is known, male hummingbirds do not take part in nesting. Most species make a cup-shaped nest on the branch of a tree or shrub. Two white eggs are laid which, despite being the smallest of all bird eggs, are in fact large relative to the hummingbird’s adult size. Incubation typically lasts 12–19 days. The nest varies in size relative to species, from smaller than half of a walnut shell to several centimeters in diameter.

Hummingbirds are found only in the Americas, from southern Alaska to Tierra del Fuego, including the Caribbean. The majority of species occur in tropical Central and South America, but several species also breed in temperate areas. Only the migratory Ruby-throated Hummingbird breeds in continental North America east of the Mississippi River and Great Lakes. The Black-chinned Hummingbird, its close relative and another migrant, is the most widespread and common species in the western United States, while the Rufous Hummingbird is the most widespread species in western Canada.

 

 

Most hummingbirds of the U.S. and Canada migrate south in fall to spend the northern winter in Mexico or Central America. A few southern South American species also move to the tropics in the southern winter. A few species are year-round residents in the warmer coastal and interior desert regions. Among these is Anna’s Hummingbird, a common resident from southern California inland to southern Arizona and north to southwestern British Columbia.